This website will bring together the results of an NSF-funded project to study the diversification of a large
clade of Apiaceae from primarily the western United States. This clade, known as the Perennial Endemic North
American (PENA) clade of Apiaceae, contains 18 genera and over 200 species. The results of this project will be
presented here in the form of an electronic monograph that will include species descriptions, photographs,
specimen records, distribution maps, phylogenetic trees, and more. This is a work in progress, and we hope to
complete it by 2024. An abstract of our project, Collaborative Research: ARTS: Diversification in the Perennial
Endemic North American (PENA) clade of Apiaceae: Defining genera and species in a major western North American
radiation, can be found here.
Despite its size and importance in the western flora, the evolution of the Perennial Endemic North American
(PENA) clade of Apiaceae is poorly understood, a fact reflected in the artificiality of its genera. This project
will develop molecular-phylogenetic hypotheses of the PENA clade using comprehensive sampling of all known species
and infraspecific taxa to serve as a framework to test specific hypotheses on species delimitation and relationships
congruent with morphology, ecology, geography, and climate. In diverse and rapidly evolving groups, such the PENA
clade, the inability to accurately define species boundaries limits any discussion of a plant’s value, use,
ecological relationships or conservation. Our work will utilize molecular, ecological, morphological, and
climatic data to clarify the taxonomy and relationships of species within the PENA clade and provide a revised
and updated classification for this group. An important part of this project is to develop this website, an online
electronic monograph, that will include interactive identification guides, as well as species pages with descriptions,
images, distribution maps, and links to specimen data. The identification tools and species-specific environmental
and ecological data provided on this website will aid in future research and in the conservation of the many rare
species found within this group.
Don Mansfield, Harold M. Tucker Herbarium, The College of Idaho
Jim Smith, Snake River Plains Herbarium, Boise State University
Because this study focuses on a large, diverse plant group in North America, it provides a wide range of prospects
for scientific outreach and training across a broad cross section of target audiences. Students of all levels will
be involved in this project. Undergraduate and graduate students from diverse backgrounds will be recruited to
take part in field and laboratory work. Field studies in Idaho will be organized for K-12 students and their
teachers, introducing them to the taxonomy, evolution, and ethnobotany of these plants. In addition, although the
primary goal of this project is to better delimit the taxonomy and species boundaries of the PENA clade, the
completion of this work will allow future studies to investigate patterns and processes of speciation, ecological
interactions, community assembly and provide a greater understanding for the diversification of western North
American Flora as well as the Biota that rely on that flora.
Mary Ann Feist, Wisconsin State Herbarium, University of Wisconsin-Madison